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How is the FROM field used?



When I compose a message in Mutt, and it passes my message on to my
MTA (sendmail), what happens with the contents of the FROM field?  I
thought that it was inserted into the smtp MAIL FROM command, but
someone on another list said this is not the case.  

If not, what actually is sent through the MAIL FROM command?

Also, is this allowed to be null, because it seems mutt won't let me
send a null FROM field.